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Traditionally, the priests blessed the people every morning after the sacrifice at the mishkan (and later at the Temple). Today, Sephardic synagogues end their service with this blessing as a benediction (Ashkenaz only recite it on Pesach, Shavu'ot, Rosh Hashanah, Yom Kippur, and Sukkot).
When the cantor is finishing the modim (thanksgiving) blessing ("Our God and God of our fathers, bless us with the threefold [priestly] blessing…") and reaches the word "kohanim," the Kohanim present are summoned to recite this blessing upon the assembled minyan (representing Israel). The congregation responds by saying, "am kedoshekha ka'amur."
According to midrash (and Jewish tradition), in the beginning only God could bless other people. His first blessing was to Adam and Eve ("Be fruitful..."). Later He blessed Noah and his sons when they left the teivah (ark). God especially blessed Abraham the tzaddik, who was given the power of blessing others (Gen. 12:3 - "Who you bless, I will bless"). Abraham did not directly bless Isaac, however, since he did not want to also bless Ishmael, so God stepped in and blessed Isaac directly (Gen. 25:11). The power of blessing others was then transferred to Isaac, who then transferred it to Jacob (Gen. 27:30), who blessed his twelve sons before he died (Gen. 48-49). In the Torah, God told Moses that the power to bless others would now come from His appointed kohanim.
All mitzvot should be performed joyfully, though if someone cannot attain this emotion, the mitzvah must still be performed. The mitzvah of birkat kohanim is unique in that if a kohen is unhappy, he must not recite the blessing (therefore a kohen who is in aveilut (mourning over the death of a parent) is required to leave during birkat kohanim).
The blessing recited before birkat kohanim is "l'varekh et amo Yisrael b'ahava" ("to bless His nation Israel with love"). The phrase "with love" is integral and is intended to represent the love of the LORD for His people. Just as God blesses us as an expression of His kindness, so to the kohen must seek to display God's love in this manner.
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